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The biological systematics and taxonomy of invertebrates as proposed by Richard C. Brusca and "Gary J." Brusca in 2003 is: a system of classification of invertebrates, as a way to classify animals without backbones.
Prokaryotesβ»
- Kingdom Eubacteria, also known as Bacteria β Domain of microorganisms
- Kingdom Archaea, also known as Archaebacteria β Domain of single-celled organisms
Eukaryotes (Eukaryota, or Eukarya)β»
- Kingdom Fungi β Biological kingdom, "separate from plants." And animals
- Kingdom Plantae β Kingdom of photosynthetic eukaryotes (= Metaphyta)
- Kingdom Protista β Eukaryotes other than animals, plants or fungi
- Phylum Euglenida β Class of protozoans
- Phylum Kinetoplastida β Flagellated protists belonging to the phylum Euglenozoa
- Alveolata phyla β Superphylum of protists
- Phylum Ciliophora β Taxon of protozoans with hair-like organelles called cilia
- Phylum Apicomplexa β Phylum of parasitic alveolates
- Phylum Dinoflagellata β Unicellular algae with two flagella
- Phylum Stramenopila β Clade of eukaryotesPages displaying short descriptions of redirect targets
- Phylum Rhizopoda β Cellular body type
- Phylum Actinopoda β Cellular body type
- Phylum Granuloreticulosa β Phylum of amoeboid protists
- Phylum Diplomonadida β Group of mostly parasitic flagellates
- Phylum Parabasilida β Group of flagellated protists
- Phylum Cryptomonada β Group of algae and colorless flagellatesPages displaying short descriptions of redirect targets
- Phylum Microspora β Taxon of autotrophic fungus-like protists
- Phylum Ascetospora β A group of eukaryotes that are parasites of animals
- Phylum Choanoflagellata β Group of eukaryotes considered the closest living relatives of animals
- Phylum Chlorophyta β Phylum of green algae
- Phylum Opalinida β Small group of peculiar heterokonts, family Opalinidae, order Slopalinida
- Incertae sedis: Genus Stephanopogon β Genus of flagellate marine protozoan
Kingdom Animalia (Metazoa)β»
Parazoaβ»
Phylum Poriferaβ»
- Phylum Porifera
- Class Calcarea
- Subclass Calcinea
- Subclass Calcaronea
- Class Hexactinellida
- Subclass Amphidiscophora
- Subclass Hexasterophora
- Class Demospongiae
- Subclass Homoscleromorpha
- Subclass Tetractinomorpha
- Subclass Ceractinomorpha
Mesozoaβ»
Phylum Placozoaβ»
- Phylum Placozoa
Phylum Monoblastozoaβ»
- Phylum Monoblastozoa
Phylum Rhombozoaβ»
- Phylum Rhombozoa
- Order Dicyemida
- Order Heterocyemida
Phylum Orthonectidaβ»
- Phylum Orthonectida
Eumetazoaβ»
Radiataβ»
Phylum Cnidariaβ»
- Phylum Cnidaria
- Class Hydrozoa
- Order Hydroida
- Suborder Anthomedusae (= Gymnoblastea or Athecata)
- Suborder Leptomedusae (= Calyptoblastea or Thecata)
- Order Trachylina
- Order Siphonophora
- Order Chondrophora
- Order Actinulida
- Class Anthozoa
- Subclass Octocorallia (= Alcyonaria)
- Order Alcyonacea
- Order Gastraxonacea
- Order Gorgonacea
- Order Helioporacea
- Order Pennatulacea
- Order Protoalcyonaria
- Order Stolonifera
- Order Telestacea
- Subclass Hexacorallia (= Zoantharia)
- Order Actiniaria
- Order Scleractinia (= Madreporaria)
- Order Zoanthidea
- Order Corallimorpharia
- Subclass Ceriantipatharia
- Order Antipatharia
- Order Ceriantharia
- Order Stauromedusae
- Order Coronatae
- Order Semaeostomae
- Order Rhizostomae
Phylum Ctenophoraβ»
- Phylum Ctenophora
- Order Beroida
- Order Cestida
- Order Cydippida
- Order Ganeshida
- Order Lobata
- Order Platyctenida
- Order Thalassocalycida
Bilateriaβ»
The authors divide the bilaterians in three informal groups:
- acoelomates (phyla Platyhelminthes, Gastrotricha, Entoprocta, Gnathostomulida, Cycliophora)
- blastocoelomate (or pseducoelomate, phyla Rotifera, Kinorhyncha, Nematoda, Nematomorpha, Acanthocephala, Loricifera)
- coelomates (or eucoelomates, phyla Nemertea, Priapula, Annelida, Sipuncula, Echiura, Onychophora, Tardigrada, Arthropoda, Mollusca, Phoronida, Ectoprocta, Brachiopoda, Echinodermata, Chaetognatha, Hemichordata, Chordata).
Several groups traditionally viewed as having blastocoelomate condition are viewed here as acoelomates (e.g., Gastrotricha, Entoprocta, Gnathostomulida).
Some of the coelomates groups (e.g., Arthropoda, Mollusca) have greatly reduced celomic spaces; often the "main body cavity is a bloodfilled space called a hemocoel." And is associated with an open circulatory system.
The Brachiopoda, Ectoprocta and Phoronida are viewed as lophophorates.
In a phylogeny, the bilaterians are divided in:
Phylum Platyhelminthesβ»
- Phylum Platyhelminthes
- Class Turbellaria
- Order Acoela
- Order Catenulida
- Order Haplopharyngida
- Order Lecithoepitheliata
- Order Macrostomida
- Order Nemertodermatida
- Order Polycladida
- Order Prolecithophora
- Order Proplicastomata
- Order Proseriata
- Order Rhabdocoela
- Suborder Dalyellioida
- Suborder Thyphloplanoida
- Suborder Kalyptorhynchia
- Suborder Temnocephalida
- Order Tricladida
- Class Monogenea
- Subclass Monopisthocotylea
- Subclass Polyopisthocotylea
- Class Trematoda
- Subclass Digenea
- Subclass Aspidogastrea
- Class Cestoda
- Subclass Cestodaria
- Subclass Eucestoda
Phylum Nemerteaβ»
- Phylum Nemertea
Phylum Rotiferaβ»
- Phylum Rotifera
- Class Digonata
- Order Seisonidea
- Order Bdelloidea
- Class Monogononta
Phylum Gastrotrichaβ»
- Phylum Gastrotricha
- Order Macrodasyida
- Order Chaetonotida
- Suborder Multitubulatina
- Suborder Paucitubulatina
Phylum Kinorhynchaβ»
- Phylum Kinorhyncha
- Order Cyclorhagida
- Order Homalorhagida
Phylum Nematoda (= Nemata)β»
- Class Adenophorea (= Aphasmida)
- Subclass Enoplia
- Subclass Chromadoria
- Class Secernentea (= Phasmida)
- Subclass Rhabditia
- Subclass Spiruria
- Subclass Diplogasteria
Phylum Nematomorphaβ»
- Phylum Nematomorpha
- Order Nectonematoidea
- Order Gordioidea
Phylum Priapulaβ»
- Phylum Priapula
- Family Priapulidae
- Family Tubiluchidae
- Family Maccabeidae (= Chaetostephanidae)
Phylum Acanthocephalaβ»
- Phylum Acanthocephala
- Class Palaeacanthocephala
- Class Archiacanthocephala
- Class Eoacanthocephala
Phylum Entoprocta (= Kamptozoa)β»
- Phylum Entoprocta (= Kamptozoa)
- Family Loxosomatidae
- Family Loxokalypodidae
- Family Pedicellinidae
- Family Barentsiidae
Phylum Gnathostomulidaβ»
- Phylum Gnathostomulida
- Order Filospermoidea
- Order Bursovaginoidea
Phylum Loriciferaβ»
- Phylum Loricifera
- Order Nanaloricida
- Family Nanaloricidae
- Family Pliciloricidae
Phylum Cycliophoraβ»
- Phylum Cycliophora
Phylum Annelidaβ»
- Phylum Annelida
- Class Polychaeta, with 25 orders and 87 families (not all are listed)
- Order Capitellida
- Order Chaetopterida
- Order Cirratulida
- Order Eunicida
- Order Myzostomida
- Order Opheliida
- Order Spionida
- Order Orbiniida
- Order Oweniida
- Order Phyllodocida
- Order Sabellida
- Order Terebellida
- Class Clitellata
- Subclass Oligochaeta
- Order Lumbriculida
- Family Lumbriculidae
- Order Moniligastrida
- Family Moniligastridae
- Order Haplotaxida, with 25 families (not all are listed)
- Family Almidae
- Family Megascolecidae
- Family Tubificidae
- Family Naididae
- Family Lumbricidae
- Subclass Hirudinoidea
- Order Acanthobdellida
- Order Branchiobdellida
- Order Hirudinida
Phylum Sipunculaβ»
- Phylum Sipuncula
- Class Phascolosomida
- Order Aspidosiphoniformes
- Order Phascolosomiformes
- Class Sipunculida
- Order Golfingiaformes
- Family Themistidae
- Family Phascolionidae
- Family Golfingiidae
- Order Sipunculiformes
Phylum Echiuraβ»
- Phylum Echiura
- Order Echiuroinea
- Order Xenopneusta
- Order Heteromyota
Phylum Onychophoraβ»
- Phylum Onychophora
- Family Peripatidae
- Family Peripatopsidae
Phylum Tardigradaβ»
- Phylum Tardigrada
- Order Heterotardigrada
- Order Mesotardigrada
- Order Eutardigrada
Phylum Arthropodaβ»
- Phylum Arthropoda
- Subphylum Trilobitomorpha
- Subphylum Crustacea
- Class Remipedia
- Class Cephalocarida
- Class Branchiopoda
- Order Anostraca
- Order Notostraca
- Order Diplostraca
- Suborder Laevicaudata
- Suborder Spinicaudata
- Suborder Cyclestherida
- Suborder Cladocera
- Class Malacostraca
- Subclass Phyllocarida
- Order Lepstostraca
- Subclass Eumalacostraca
- Superorder Hoplocarida
- Order Stomatopoda
- Superorder Syncarida
- Order Bathynellacea
- Order Anaspidacea
- Superorder Eucarida
- Order Euphausiacea
- Order Amphionidacea
- Order Decapoda
- Suborder Dendrobranchiata
- Suborder Pleocyemata
- Infraorder Caridea
- Infraorder Stenopodidea
- Infraorder Brachyura
- Infraorder Anomura
- Infraorder Astacidea
- Infraorder Palinura
- Infraorder Thalassinidea
- Superorder Peracarida
- Order Mysida
- Order Lophogastrida
- Order Cumacea
- Order Tanaidacea
- Order Mictacea
- Order Spelaeogriphacea
- Order Thermosbaenacea
- Order Isopoda
- Suborder Anthuridea
- Suborder Asellota
- Suborder Calabozoidea
- Suborder Epicaridea
- Suborder Flabellifera
- Suborder Gnathiidea
- Suborder Oniscidea
- Suborder Phreatoicidea
- Suborder Valvifera
- Order Amphipoda
- Suborder Gammaridea
- Suborder Hyperiidea
- Suborder Caprellidea
- Suborder Ingolfiellidea
- Class Maxillopoda
- Subclass Thecostraca
- Infraclass Facetotecta
- Infraclass Ascothoracida
- Infraclass Cirripedia
- Superorder Acrothoracica
- Superorder Rhizocephala
- Superorder Thoracica
- Subclass Tantulocarida
- Subclass Branchiura
- Subclass Pentastomida
- Subclass Mystacocarida
- Subclass Copepoda
- Infraclass Progymnoplea
- Order Platycopioida
- Infraclass Neocopepoda
- Order Calanoida
- Order Cyclopoida
- Order Gelyelloida
- Order Harpacticoida
- Order Misophrioida
- Order Monstrilloida
- Order Mormonilloida
- Order Poecilostomatoida
- Order Siphonostomatoida
- Subclass Ostracoda
- Superorder Myodocopa
- Order Myodocopida
- Order Halocyprida
- Superorder Podocopa
- Order Podocopida
- Order Platycopida
- Order Palaeocopida
- Subphylum Hexapoda
- Class Entognatha
- Order Collembola
- Order Protura
- Order Diplura
- Class Insecta
- Subclass Archaeognatha
- Order Archaeognatha (= Microcoryphia)
- Subclass Zygentoma
- Order Thysanura
- Subclass Pterygota
- Infraclass Palaeoptera
- Order Ephemeroptera
- Order Odonata
- Infraclass Neoptera
- Order Plecoptera
- Order Blattodea
- Order Isoptera
- Order Mantodea
- Order Phasmida (= Phasmatoptera)
- Order Grylloblattodea
- Order Dermaptera
- Order Orthoptera
- Order Mantophasmatodea
- Order Embioptera
- Order Zoraptera
- Order Psocoptera
- Order Phthiraptera
- Order Thysanoptera
- Order Hemiptera
- Order Strepsiptera
- Order Megaloptera
- Order Raphidioptera
- Order Neuroptera
- Order Coleoptera
- Order Mecoptera
- Order Siphonaptera
- Order Diptera
- Order Trichoptera
- Order Lepidoptera
- Order Hymenoptera
- Subphylum Myriapoda
- Class Diplopoda
- Subclass Penicillata
- Order Polyxenida
- Subclass Chilognatha
- Order Callipodida
- Order Chordeumatida
- Order Glomerida
- Order Glomeridesmida
- Order Julida
- Order Platydesmida
- Order Polydesmida
- Order Polyzoniida
- Order Siphonophorida
- Order Siphoniulida
- Order Sphaerotheriida
- Order Spirobolida
- Order Spirostreptida
- Order Stemmiulida
- Class Chilopoda
- Subclass Notostigmophora
- Order Scutigeromorpha
- Subclass Pleurostigmophora
- Order Craterostigmomorpha
- Order Geophilomorpha
- Order Lithobiomorpha
- Order Scolopendromorpha
- Subphylum Cheliceriformes
- Class Pycnogonida
- Class Chelicerata
- Subclass Merostomata
- Order Eurypterida
- Order Xiphosura
- Subclass Arachnida
- Order Acari
- Suborder Opiloacarifomes
- Suborder Parasitiformes
- Suborder Acariformes
- Suborder Mesothelae
- Suborder Opisthothelae
- Superfamily Mygalomorpha
- Family Ctenizidae
- Family Atypidae
- Family Theraphosidae
- Family Dipluridae
- Superfamily Araneomorphae
- Family Loxoscelidae
- Family Theridiidae
- Family Uloboridae
- Family Araneidae
- Family Tetragnathidae
- Family Clubionidae
- Family Linyphiidae
- Family Agelenidae
- Family Argyronetidae
- Family Lycosidae
- Family Pisauridae
- Family Oxyopidae
- Family Thomisidae
- Family Heteropodidae
- Family Salticidae
- Family Dinopidae
- Family Scytodidae
- Order Opiliones
- Order Palpigradi
- Order Pseudoscorpionida
- Order Ricinulei
- Order Schizomida
- Order Scorpiones
- Order Solpugida
- Order Uropygi
Phylum Molluscaβ»
- Phylum Mollusca
- Class Aplacophora
- Subclass Chaetodermomorpha (= Caudofoveata)
- Subclass Neomeniomorpha (= Solenogastres)
- Class Monoplacophora
- Class Polyplacophora
- Order Lepidopleurida
- Order Ischnochitonida
- Order Acanthochitonida
- Class Gastropoda
- Subclass Prosobranchia
- Order Archaeogastropoda
- Order Mesogastropoda
- Order Neogastropoda
- Subclass Opisthobranchia
- Order Acochlidioidea
- Order Cephalaspidea
- Order Runcinoidea
- Order Sacoglossa
- Order Anaspidea
- Order Thecosomata
- Order Gymnosomata
- Order Notaspidea
- Order Nudibranchia
- Subclass Pulmonata
- Order Archaeopulmonata
- Order Basommatophora
- Order Stylommatophora
- Order Systellommatophora
- Class Bivalvia (=Pelecypoda, or Lamellibranchiata)
- Subclass Protobranchia
- Order Nuculida (= Palaeotaxodonta)
- Order Solemyida (= Cryptodonta)
- Subclass Lamellibranchia
- Superorder Filibranchia (= Pteriomorpha)
- Superorder Eulamellibranchia (= Heterodonta)
- Order Palaeoheterodonta
- Order Veneroida
- Order Myoida
- Subclass Anomalodesmata
- Class Scaphopoda
- Class Cephalopoda (= Siphonopoda)
- Subclass Nautiloidea (= Tetrabranchiata)
- Subclass Coleoidea (= Dibranchiata)
- Order Sepioida
- Order Teuthoida (= Decapoda)
- Order Octopoda
- Order Vampyromorpha
Phylum Phoronidaβ»
- Phylum Phoronida
Phylum Ectoprocta (= Bryozoa)β»
- Phylum Ectoprocta (= Bryozoa)
- Class Phylactolaemata
- Class Stenolaemata
- Class Gymnolaemata
- Order Ctenostomata
- Order Cheilostomata
Phylum Brachiopodaβ»
- Phylum Brachiopoda
- Class Inarticulata
- Class Articulata
Phylum Echinodermataβ»
- Phylum Echinodermata
- Class Crinoidea
- Class Asteroidea
- Order Platysterida
- Order Paxillosida
- Order Valvatida
- Order Spinulosida
- Order Forcipulatida
- βSea daisiesβ (previously the class Concentricycloidea, but assigned by many authorities to the Spinulosida)
- Class Ophiuroidea
- Order Oegophiurida
- Order Phrynophiurida
- Order Ophiurida
- Class Echinoidea
- Subclass Cidaroidea
- Subclass Euechinoidea
- Infraclass Echinothurioidea
- Infraclass Acroechinoidea
- Cohort Diadematacea
- Cohort Echinacea
- Cohort Irregularia
- Class Holothuroidea
- Subclass Dendrochirotacea
- Order Dactylochirotida
- Order Dendrochirotida
- Subclass Aspidochirotacea
- Order Aspidochirotida
- Order Elasipodida
- Subclass Apodacea
Phylum Chaetognathaβ»
- Phylum Chaetognatha
- Order Phragmophora
- Order Aphragmophora
Phylum Hemichordataβ»
- Phylum Hemichordata
- Class Enteropneusta
- Class Pterobranchia
- Class Planctosphaeroidea
Phylum Chordataβ»
- Phylum Chordata
- Subphylum Urochordata (= Tunicata)
- Class Ascidiacea
- Class Thaliacea
- Class Appendicularia (= Larvacea)
- Class Sorberacea
- Subphylum Cephalochordata (= Acrania)
- Subphylum Vertebrata
- Class Myxini
- Class Cephalaspidomorphi
- Class Chondrichthyes
- Class Osteichthyes
- Class Amphibia
- Class Reptiliomorpha (= Sauropsida)
- Class Mammalia
See alsoβ»
- Taxonomy of the Gastropoda (Bouchet et al., 2017)
- Taxonomy of the Gastropoda (Bouchet & Rocroi, 2005) β System for the scientific classification of gastropod mollusks
- Taxonomy of the Gastropoda (Ponder & Lindberg, 1997) β Morphological system for the scientific classification of gastropod mollusks
- Taxonomy of the Conoidea (Tucker & Tenorio, 2009) β Classification of cone snails and allies
- Taxonomy of the Lepidoptera β Classification of moths and butterflies
Referencesβ»
- ^ Richard C. Brusca & Gary J. Brusca (2003). Invertebrates (2nd ed.). Sunderland, Massachusetts: Sinauer Associates. ISBN 978-0-87893-097-5.
- ^ Brusca, R. C.; Brusca, G. J. (2005). Invertebrados (2nd ed.). Madrid: McGraw-Hill-Interamericana. ISBN 978-0-87893-097-5.
- ^ Richard C. Brusca & Gary J. Brusca (2003), p. 875.